1. Institute of Tibet Plateau Ecology, Tibet Agricultural & Animal Husbandry University, Nyingchi 860000, China
2. Key Laboratory of Forest Ecology in Tibet Plateau, Tibet Agricultural & Animal Husbandry University, Ministry of Education, Nyingchi 860000, China
3. National Forest Ecosystem Observation & Research Station of Nyingchi Tibet, Nyingchi 860000, China
4. Key Laboratory of Alpine Vegetation Ecological Security in Tibet, Nyingchi 860000, China