1. Hubei International Scientific and Technological Center of Ecological Conservation and Management in the Three Gorges Area, Engineering Research Center of Eco-Environment in Three Gorges Reservoir Region, Ministry of Education, College of Biological and Pharmaceutical Sciences, China Three Gorges University, Yichang 443002, China
2. Hubei Key Laboratory of Rare Resource Plants in Three Gorges Reservoir Area, China Three Gorges Corporation, Yichang 443000, China
3. College of Hydraulic and Environmental Engineering, China Three Gorges University, Yichang 443002, China