1. Department of Botany and Zoology, Biosciences Center, Federal University of Rio Grande do Norte (UFRN), Highway BR-101, Lagoa Nova, Campus Universitario, Natal 59072-970, RN, Brazil
2. Institute of Systematics and Evolution of Animals, Polish Academy of Sciences, Sławkowska 17, Pl-31-016 Kraków, Poland
3. Institut National De La Recherche Agronomique Clermont-Auvergne-Rhônes-Alpe, INRAE, 63000 Clermont-Ferrand, France