1. RIKEN Center for Sustainable Resource Science, 1-7-22, Suehiro-cho, Tsurumi-ku, Yokohama 230-0045, Kanagawa, Japan
2. Research Center for Subtropical Fisheries, Seikai National Fisheries Research Institute, Japan Fishery Research and Education Agency, 148 Fukaiota, Ishigaki 907-0451, Okinawa, Japan
3. Graduate School of Medical Life Science, Yokohama City University, 1-7-29 Suehiro-cho, Tsurumi-ku, Yokohama 230-0045, Kanagawa, Japan
4. Graduate School of Bioagricultural Sciences, Nagoya University, 1 Furo-cho, Chikusa-ku, Nagoya 464-8601, Aichi, Japan