1. Faculty of Sciences of Bizerte, University of Carthage, Jarzouna 7021, Tunisia
2. Laboratory of Forest Ecology (LR11INRGREF03), National Institute of Research in Rural Engineering, Water and Forests (INRGREF), University of Carthage, Hédi Elkarray Street, Elmenzah IV, BP 10, Ariana 2080, Tunisia
3. Direction de la Recherche Forestière, Ministère des Ressources Naturelles et des Forêts, 2700, Rue Einstein, Quebec, QC G1P 3W8, Canada
4. Centre for Forest Research and Institute for Systems and Integrative Biology, Université Laval, 1030 Avenue de la Médecine, Quebec, QC G1V 0A6, Canada