1. State Key Laboratory of Resource Insects, Institute of Apicultural Research, Chinese Academy of Agricultural Sciences, Beijing 100093, China
2. Key Laboratory for Insect-Pollinator Biology of the Ministry of Agriculture and Rural Affairs, Institute of Apicultural Research, Chinese Academy of Agricultural Sciences, Beijing 100093, China
3. Department of Ecology, School of Resources and Engineering, Anhui University, Hefei 230601, China
4. Anhui Province Key Laboratory of Wetland Ecosystem Protection and Restoration, Hefei 230601, China