1. Yunnan Academy of Biodiversity, Southwest Forestry University, Kunming 650224, China
2. Yunnan Key Laboratory of Plateau Wetland Conservation, Restoration and Ecological Services, Southwest Forestry University, Kunming 650224, China
3. School of Life Science, Southwest Forestry University, Kunming 650224, China
4. School of Ecology and Environment, Yunnan University, Kunming 650091, China