1. Department of Biology and Geology, University of South Carolina-Aiken, Aiken, SC 29801, USA
2. Department of Microbiology, University of Tennessee Knoxville, Knoxville, TN 37996, USA
3. Department of Medicine and Vascular Biology Center, Medical College of Georgia, Augusta University, Augusta, GA 30912, USA
4. Arthropod-Borne Animal Diseases Research Unit, Center for Grain and Animal Health Research, Agricultural Research Service, United States Department of Agriculture, Manhattan, KS 66502, USA