1. School of Life Sciences, Key Laboratory of Conservation and Application in Biodiversity of South China, Guangzhou University, Guangzhou Higher Education Mega Center, #230 Waihuanxi Road, Guangzhou 510006, China
2. School of Earth Sciences and Engineering, Guangdong Provincial Key Lab of Geological Processes and Mineral Resources, Sun Yat-sen University, Guangzhou 510275, China
3. College of Life Sciences and Academy for Multidisciplinary Studies, Capital Normal University, Xisanhuanbeilu 105, Haidian District, Beijing 100048, China