1. School of Life Science, Qinghai Normal University, Xining 810008, China
2. Sanjiangyuan Grassland Ecosystem National Observation and Research Station, Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences, Xining 810008, China
3. Key Laboratory of Medicinal Animal and Plant Resources of Qinghai-Tibetan Plateau in Qinghai Province, Xining 810008, China
4. Academy of Plateau Science and Sustainability, People’s Government of Qinghai Province and Beijing Normal University, Xining 810016, China