1. Papanin Institute for Biology of Inland Waters, Russian Academy of Sciences, 152742 Borok, Russia
2. A.N. Severtsov Institute of Ecology and Evolution of the Russian Academy of Sciences, 119071 Moscow, Russia
3. N. Laverov Federal Center for Integrated Arctic Research of the Ural Branch of the Russian Academy of Sciences, Nikolsky Avenue 20, 163020 Arkhangelsk, Russia