1. School of Life and Sciences, Qingdao University, Qingdao 266071, China
2. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
3. College of Earth Science, University of Chinese Academy of Sciences, Beijing 100049, China
4. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao 266237, China
5. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China