1. Department of Ophthalmology, Medical College of Georgia at Augusta University, Augusta, GA 30912, USA
2. Department of Physiology, Medical College of Georgia at Augusta University, Augusta, GA 30912, USA
3. Charlie Norwood VA Medical Center, Augusta, GA 30904, USA
4. Department of Cellular Biology and Anatomy, Medical College of Georgia at Augusta University, Augusta, GA 30912, USA
5. James and Jean Culver Vision Discovery Institute, Medical College of Georgia at Augusta University, Augusta, GA 30912, USA