1. Biological and Environmental Sciences, The University of Stirling, Stirling FK9 4LA, UK
2. Geography Department, Swansea University, Singleton Park, Swansea SA2 8PP, UK
3. Permian Global Research Ltd., London WC2R 0BU, UK
4. Forest Research, Agency of the British Forestry Commission, Roslin, Midlothian, Edinburgh EH25 9SY, UK