1. Key Laboratory of South China Agricultural Plant Molecular Analysis and Genetic Improvement & Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. South China National Botanical Garden, Guangzhou 510650, China
4. CAS Engineering Laboratory for Vegetation Ecosystem Restoration on Islands and Coastal Zones, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China
5. College of Traditional Chinese Medicine, Guangdong Pharmaceutical University, Guangzhou 510006, China