1. Department of Biodiversity Ecology and Evolution, Faculty of Biological Sciences, Complutense University of Madrid, 28040 Madrid, Spain
2. Natural History Museum, London SW7 5BD, UK
3. Herpetology Lab, Natural History Museum, London SW7 5BD, UK
4. bDIV, Biology Department, Vrije Universiteit Brussel, Pleinlaan 2, 1050 Elsene, Belgium