1. Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment, Nanjing 210042, China
2. Hulun Lake Wetland Ecological Environment Scientific Observation and Research Station, Hulunbuir 021008, China
3. Hulunbuir Academy of Inland Lakes in Northern Cold & Arid Areas, Hulunbuir 021008, China
4. School of Geographic Science, Nantong University, Nantong 226007, China