1. Queen Mother Hospital for Animals, The Royal Veterinary College, Hawkshead Lane, Hatfield AL9 7TA, Hertfordshire, UK
2. Department of Clinical Science and Services, The Royal Veterinary College, Hawkshead Lane, Hatfield AL9 7TA, Hertfordshire, UK
3. Department of Pathobiology and Population Sciences, The Royal Veterinary College, Hawkshead Lane, Hatfield AL9 7TA, Hertfordshire, UK