1. Key Laboratory for Conserving Wildlife with Small Populations in Yunnan, Southwest Forestry University, Kunming 650224, China
2. Department of Biodiversity Conservation, Southwest Forestry University, Kunming 650224, China
3. Administration of Zixi Mountain Provincial Nature Reserve, Chuxiong 675008, China
4. Research Center for Biodiversity Conservation and Biosafety/State Environmental Protection Key Laboratory on Biosafety, Nanjing Institute of Environmental Sciences, Ministry of Ecology and Environment of China, Nanjing 210042, China