1. National Aquatic Resources Research and Development Agency, Crow Island, Colombo 01500, Sri Lanka
2. Department of Biology, Faculty of Sciences, Ghent University, 9000 Ghent, Belgium
3. GiMaRIS, Marine Research, Inventory & Strategy Solutions, Rijksstraatweg 75, 2171 AK Sassenheim, The Netherlands
4. Netherlands Centre for Biodiversity Naturalis, 2333 CR Leiden, The Netherlands