1. Key Laboratory of Cultivation and Protection for Non-Wood Forest Trees of Ministry of Education, Central South University of Forestry and Technology, Changsha 410004, China
2. Key Laboratory of Non-Wood Forest Products of State Forestry Administration, Central South University of Forestry and Technology, Changsha 410004, China
3. Anhui Hongsen Hi Tech Forestry Co., Ltd., Bozhou 236814, China
4. Department of Biology, Eastern New Mexico University, Portales, NM 88130, USA