1. Graduate School of Bioscience and Biotechnology, Fukui Prefectural University, Fukui 910-1195, Japan
2. Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University, Fukuoka 819-0395, Japan
3. Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo 113-8567, Japan
4. Graduate School of Agriculture, Ibaraki University, Ibaraki 300-0393, Japan