1. College of Resources and Environmental Sciences, China Agricultural University, Beijing 100193, China
2. Beijing Key Laboratory of Biodiversity and Organic Farming, China Agricultural University, Beijing 100193, China
3. Resources & Environment College, Tibet Agriculture & Animal Husbandry University, Nyingchi 860000, China
4. Key Laboratory of Forest Ecology and Management, Institute of Applied Ecology, Chinese Academy of Sciences, Shenyang 110016, China
5. Institute of Tibet Plateau Ecology, Tibet Agriculture & Animal Husbandry University, Nyingchi 860000, China
6. Key Laboratory of Forest Ecology in Tibet Plateau, Tibet Agriculture & Animal Husbandry University, Ministry of Education, Nyingchi 860000, China