1. State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences, Wuhan 430072, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Key Laboratory of Aquatic Organism Protection and Ecological Restoration in Cold Waters, Heilongjiang River Fisheries Research Institute, Chinese Academy of Fishery Sciences, Harbin 150010, China