1. Hubei Hongshan Laboratory, National Key Laboratory of Crop Genetic Improvement, Huazhong Agricultural University, Wuhan 430070, China
2. National Key Laboratory of Wheat Breeding, College of Life Sciences, Shandong Agricultural University, Taian 271018, China
3. Zhejiang Academy of Agricultural Sciences, Institute of Maize and Featured Upland Crops, Hangzhou 310015, China