1. Friedrich-Loeffler-Institut (FLI), Federal Research Institute for Animal Health, Südufer 10, 17493 Greifswald, Germany
2. Leibniz Centre for Agricultural Landscape Research (ZALF), Eberswalder Str. 84, 15374 Müncheberg, Germany
3. Institutes of Bacteriology and Parasitology, Medical Faculty, University of Strasbourg, UR 3073 PHAVI, 67000 Strasbourg, France