1. Institute for Biodiversity and Environmental Research, Universiti Brunei Darussalam, Bandar Seri Begawan BE 1410, Brunei
2. Environmental and Life Sciences Programme, Faculty of Science, Universiti Brunei Darussalam, Bandar Seri Begawan BE 1410, Brunei
3. School of Biological Sciences, University of Aberdeen Cruickshank Building, Aberdeen AB24 3UL, UK