1. CNRS, EthoS (Éthologie Animale et Humaine)—UMR 6552, Univ Rennes, Normandie Univ, F-35000 Rennes, France
2. CNRS, Université de Paris Cité, UMR 8002, Integrative Neuroscience and Cognitive Center, F-75270 Paris, France
3. Department Zoology and Entomology, Rhodes University, Makhanda 6139, South Africa