1. The UWA Institute of Agriculture, The University of Western Australia, Perth, WA 6009, Australia
2. Centre for Plant Sciences, Faculty of Biology, University of Leeds, Leeds LS2 9JT, UK
3. Food and Plant Biology Group, Departamento de Biología Vegetal, Facultad de Agronomía, Universidad de la República, Sayago CP 12900, Uruguay
4. School of Biosciences, College of Life and Environmental Sciences, University of Birmingham, Edgbaston B15 2TT, UK
5. Horticulture and Irrigated Agriculture, Department of Primary Industries and Regional Development, South Perth, WA 6151, Australia