1. Biology Department, College of Natural and Computational Sciences, Hawassa University, Hawassa P.O. Box 05, Ethiopia
2. School of Plant and Horticultural Sciences, College of Agriculture, Hawassa University, Hawassa P.O. Box 05, Ethiopia
3. Institute of Soil Science and Land Evaluation, Biogeophysics, University of Hohenheim, 70593 Stuttgart, Germany
4. Quantitative Biodiversity Dynamics (QBD), Department of Biology, University of Utrecht, Padualaan 8, 3584 CH Utrecht, The Netherlands