1. Department of Integrative Biology and Biodiversity Research, Institute of Zoology, University of Natural Resources and Life Sciences Vienna (BOKU), Gregor Mendel Straße 33, 1180 Vienna, Austria
2. Department of Landscape, Spatial and Infrastructure Science, Institute of Statistics, University of Natural Resources and Life Sciences Vienna (BOKU), Peter-Jordan-Straße 82, 1190 Vienna, Austria
3. Agro-Environmental Research Centre, Institute of Environmental Sciences, Hungarian University of Agriculture and Life Sciences, Herman Ottó út 15, H-1022 Budapest, Hungary