1. State Key Laboratory of Swine and Poultry Breeding Industry Guangdong Key Laboratory of Animal Breeding and Nutrition Institute of Animal Science, Guangdong Academy of Agricultural Sciences, Guangzhou 510640, China
2. Department of Biotechnology, College of Natural and Computational Science, Aksum University, Aksum 1010, Tigray, Ethiopia
3. Department of Animal Science, Adigrat University, Adigrat 7040, Tigray, Ethiopia
4. Agricultural Science and Engineering School, Liaocheng University, Liaocheng 252000, China