1. Department of Microbiology and Biochemistry of Dairy Products, Dairy Research Institute of Asturias (IPLA), Spanish National Research Council (CSIC), Paseo Río Linares S/N, 33300 Villaviciosa, Asturias, Spain
2. Health Research Institute of Asturias (ISPA), 33011 Oviedo, Asturias, Spain
3. Laboratory of Technological Ecophysiology, Reference Centre for Lactobacilli (CERELA-CONICET), Chacabuco 145, San Miguel de Tucumán 4000, Tucumán, Argentina
4. Department of Food Microbiology, University of San Pablo Tucumán, Av. Solano Vera y Camino a Villa Nougués, San Pablo 4129, Tucumán, Argentina