1. State Key Laboratory of Cryospheric Sciences, Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China
2. University of Chinese Academy of Sciences, No. 19A Yuquan Road, Beijing 100049, China
3. Key Laboratory of Extreme Environmental Microbial Resources and Engineering, Lanzhou 730000, China
4. Key Laboratory of Desert and Desertification, Northwest Institute of Eco-Environment and Resources, Chinese Academy of Sciences, Lanzhou 730000, China
5. School of Biological and Pharmaceutical Engineering, Lanzhou Jiaotong University, Lanzhou 730070, China