1. Hainan Academy of Ocean and Fisheries Sciences, Haikou 571126, China
2. Yazhou Bay Innovation Institute, Hainan Tropical Ocean University, Sanya 572025, China
3. School of Marine Science and Technology, Tianjin University, Tianjin 300072, China
4. Ocean and Ecosystem Sciences Division, Fisheries and Oceans Canada, Bedford Institute of Oceanography, Dartmouth, NS B2Y4A2, Canada
5. College of Marine and Environmental Science, Tianjin University of Science and Technology, Tianjin 300457, China