1. School of Life and Health, Hainan University, Haikou 570228, China
2. Key Laboratory of Biology and Genetic Resources of Tropical Crops, Ministry of Agriculture and Rural Affairs, Institute of Tropical Bioscience and Biotechnology, Chinese Academy of Tropical Agricultural Sciences, Haikou 571101, China
3. Hainan Academy of Tropical Agricultural Resource, Haikou 571101, China
4. Rubber Research Institute, Chinese Academy of Tropical Agricultural Sciences, Danzhou 571737, China