1. Key Laboratory of Tropical Aquatic Germplasm of Hainan Province, Sanya Oceanographic Institution, Ocean University of China, Sanya 572000, China
2. Southern Marine Science and Engineer Guangdong Laboratory, Guangzhou 511458, China
3. MOE Key Laboratory of Marine Genetics and Breeding, Ocean University of China, Qingdao 266003, China
4. Institute of Evolution & Marine Biodiversity, Ocean University of China, Qingdao 266003, China