1. Laboratory of Biotechnology and Biomonitoring of the Environment and Oasis Ecosystems, Faculty of Sciences of Gafsa, Gafsa 2112, Tunisia
2. Department of Biology, College of Sciences, Imam Mohammad Ibn Saud Islamic University (IMSIU), Riyadh 11623, Saudi Arabia
3. Department of Zoology, College of Science, King Saud University, Riyadh 11451, Saudi Arabia
4. Institut Supérieur des Sciences et Technologies de l’Environnement Borj Cédria, Université de Carthage, Hammam chat 2050, Ben Arous, Tunis 1073, Tunisia