1. Key Laboratory of Animal Genetics, Breeding and Reproduction of Ministry of Agriculture and Rural Affairs, Institute of Animal Science, Chinese Academy of Agricultural Sciences, Beijing 100193, China
2. Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, Beijing 100101, China
3. Small Ruminant Genomics, International Center for Agricultural Research in the Dry Areas (ICARDA), Addis Ababa P.O. Box 5689, Ethiopia
4. Animal and Veterinary Sciences, SRUC and Centre for Tropical Livestock Genetics and Health (CTLGH), The Roslin Institute Building, Midlothian EH25 9RG, UK