1. School of Agriculture and Animal Science, Obihiro University of Agriculture and Veterinary Medicine, Obihiro 080-8555, Hokkaido, Japan
2. Department of Biochemistry, Asahikawa Medical University, Asahikawa 078-8510, Hokkaido, Japan
3. Laboratory for Molecular and Developmental Biology, Institute for Protein Research, Osaka University, Suita 565-0871, Osaka, Japan