1. State Key Laboratory of Continental Dynamics, Shaanxi Key Laboratory of Early Life & Environments, Department of Geology, Northwest University, Xi’an 710069, China
2. Biodiversity Institute and Department of Ecology & Evolutionary Biology, University of Kansas, Lawrence, KS 66045, USA
3. Department of Palaeontology, University of Vienna, Althanstrasse 14, 1090 Vienna, Austria
4. Palaeobiology, Department of Earth Sciences, Uppsala University, SE-752 36 Uppsala, Sweden