1. Department of Life and Food Sciences, Obihiro University of Agriculture and Veterinary Medicine, Obihiro 080-8555, Hokkaido, Japan
2. Research Center for Global Agromedicine, Obihiro University of Agriculture and Veterinary Medicine, Obihiro 080-8555, Hokkaido, Japan
3. Volunteer Bander, Division of Avian Conservation, Yamashina Institute for Ornithology, Abiko 270-1145, Chiba, Japan