1. CAS and Shandong Province Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. School of Marine Science and Engineering, Qingdao Agricultural University, Qingdao 266109, China
5. The Innovation of Seed Design, Chinese Academy of Sciences, Wuhan 430072, China