1. Yantai Institute of Coastal Zone Research, Chinese Academy of Sciences, Yantai 264003, China
2. University of Chinese Academy of Sciences, Beijing 100049, China
3. Marine Mammal and Marine Bioacoustics Laboratory, Institute of Deep-Sea Science and Engineering, Chinese Academy of Sciences, Sanya 572000, China
4. Institute of Zoology, Zoological Society of London, London NW1 4RY, UK