1. Center for Molecular Medicine, MaineHealth Institute for Research, Scarborough, ME 04074, USA
2. Department of Anatomy, Cell Biology and Physiology, Orthopaedic Surgery, School of Medicine, Indiana University, Indianapolis, IN 46202, USA
3. Indiana Center for Musculoskeletal Health, Indiana University, Indianapolis, IN 46202, USA
4. Department of Cancer Biology, Dana-Farber Cancer Institute, Harvard Medical School, Boston, MA 02115, USA
5. Department of Cell Biology, Harvard Medical School, Boston, MA 02115, USA