1. College of Forestry, Guizhou University, Guiyang 550025, China
2. Forest Ecology Research Center, Guizhou University, Guiyang 550025, China
3. Qingyuan Forest, National Observation and Research Station, Shenyang 110016, China
4. Guizhou Libo Observation and Research Station for Karst Forest Ecosystem, National Forestry and Grassland Administration, Guiyang 550025, China