1. Laboratory Ecology, Systematics, and Conservation of Biodiversity (LESCB), URL-CNRST N°18, FS, Abdelmalek Essaadi University, Tetouan 931000, Morocco
2. National Institute of Agricultural Research, Regional Center of Agricultural Research, Avenue Ennasr, BP 415 Rabat Principale, Rabat 10090, Morocco
3. Dipartimento di Geoscienze, Università degli Studi di Padova, 35131 Padova, Italy