1. College of Tropical Crops, Hainan University, Haikou 570228, China
2. Key Laboratory of Biology and Genetic Resources of Tropical Crops, Ministry of Agriculture, Hainan Institute for Tropical Agricultural Resources, Institute of Tropical Bioscience and Biotechnology, Chinese Academy of Tropical Agricultural Sciences, Haikou, 571101, China
3. College of Horticulture, China Agricultural University, Beijing 100083, China