1. CAS Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture, Wuhan Botanical Garden, The Innovative Academy of Seed Design, Chinese Academy of Sciences, Wuhan 430074, China
2. Hubei Hongshan Laboratory, Wuhan 430070, China
3. University of Chinese Academy of Sciences, Beijing 100049, China
4. Sino-African Joint Research Center, Chinese Academy of Sciences, Wuhan 430074, China