1. Jiangsu Key Laboratory of Marine Bioresources and Eco-Environment, Jiangsu Ocean University, Lianyungang 222005, China
2. Jiangsu Institute of Marine Resources Development, Lianyungang 222005, China
3. A Co-Innovation Center of Jiangsu Marine Bio-Industry Technology, Lianyungang 222005, China
4. Jiangsu Key Laboratory of Marine Biotechnology, Jiangsu Ocean University, Lianyungang 222005, China